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Graduate College |
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Courses |
BiosciencesDirector: Andrew F. RussoAffiliated faculty: Paul Abbas (Speech Pathology and Audiology/Otolaryngology--Head and Neck Surgery), Francois Abboud (Internal Medicine), Harold Adams (Neurology), Lee-Ann Allen (Internal Medicine), Michael Anderson (Molecular Physiology and Biophysics), Steven Anderson (Neurology), Nancy Andreasen (Psychiatry), Michael Apicella (Microbiology), Mark Arnold (Chemistry), Nikolai Artemyev (Molecular Physiology and Biophysics), Mario Ascoli (Pharmacology), Robert Ashman (Internal Medicine), Zuhair Ballas (Internal Medicine), Botond Banfi (Anatomy and Cell Biology), Christopher Benson (Internal Medicine), Ramesh Bhalla (Anatomy and Cell Biology), Debashish Bhattacharya (Biological Sciences), Jackie Bickenbach (Anatomy and Cell Biology), Gail Bishop (Microbiology), Daniel Bonthius (Pediatrics), Terry Braun (Biomedical Engineering), Timothy Brennan (Anesthesiology), Garry Buettner (Radiation Oncology), John Butler (Microbiology), Kevin Campbell (Molecular Physiology and Biophysics), Donald Cannon (Chemistry), Thomas Casavant (Electrical and Computer Engineering), Martin Cassell (Anatomy and Cell Biology), Mark Chapleau (Internal Medicine), Chi-Lien Cheng (Biological Sciences), Steven Clegg (Microbiology), Michael Cohen (Pathology), Kelly James Cole (Integrative Physiology), John Colgan (Internal Medicine), Josep Comeron (Biological Sciences), Robert Cornell (Anatomy and Cell Biology), Charles Cox (Microbiology), Joseph Cullen (Surgery), Michael Dailey (Biological Sciences), Morris Dailey (Pathology), Warren Darling (Integrative Physiology), Beverly Davidson (Internal Medicine), Deborah Dawson (Preventive and Community Dentistry), Kris DeMali (Biochemistry), Jeffrey Denburg (Biological Sciences), Natalie Denbury (Neurology), Frederick Domann (Radiation Oncology), John Donelson (Biochemistry), Jonathan Doorn (Pharmacy), Ken Dornfeld (Radiation Oncology), Michael Duffel (Pharmacy), Adam Dupuy (Anatomy and Cell Biology), Daniel Eberl (Biological Sciences), Adrian Elcock (Biochemistry), Hatem El-Shanti (Pediatrics), John Engelhardt (Anatomy and Cell Biology), Sarah England (Molecular Physiology and Biophysics), Frank Faraci (Internal Medicine), Jan Fassler (Biological Sciences), Michael Feiss (Microbiology), Robert Felder (Internal Medicine), Elizabeth Field (Internal Medicine), R. William Field (Occupational and Environmental Health), Rory Fisher (Pharmacology), Robert Franciscus (Anthropology), Joseph Frankel (Biological Sciences), Sonya Franklin (Chemistry), John Freeman (Psychology), George Friestad (Chemistry), Ernesto Fuentes (Biochemistry), Laurence Fuortes (Occupational and Environmental Health), Minnetta Gardinier (Pharmacology), Lei Geng (Chemistry), Frederic Gerr (Occupational and Environmental Health), Pamela Geyer (Biochemistry), James Gloer (Chemistry), Harold Goff (Chemistry), Pedro Gonzalez-Alegre (Neurology), Jean Gordon (Speech Pathology and Audiology), Prabhat Goswami (Radiation Oncology), Thomas Grabowski (Neurology), Steven Green (Biological Sciences), Thomas Griffith (Urology), Gary Gussin (Biological Sciences), Lilach Hadany (Biological Sciences), Amanda Haes (Chemistry), Gregory Hageman (Ophthalmology), Donna Hammond (Anesthesiology), John Harty (Microbiology), Eliot (Richard) Hazeltine (Psychology), Paul Heidger (Anatomy and Cell Biology), Donald Heistad (Internal Medicine), Johannes Hell (Pharmacology), Stephen Hendrix (Biological Sciences), Michael Henry (Molecular Physiology and Biophysics), Jonathan Heusel (Pathology), Raymond Hohl (Internal Medicine), Keri Hornbuckle (Civil and Environmental Engineering), Mary Horne (Pharmacology), Alexander Horswill (Microbiology), Diana Horton (Biological Sciences), Bruce Hostager (Pediatrics), Douglas Houston (Biological Sciences), Jon Houtman (Microbiology), Matthew Howard (Neurosurgery), Richard Hurtig (Speech Pathology and Audiology), Fiorenza Ianzini (Radiology), Erin Irish (Biological Sciences), Toshiki Itoh (Pathology), Jean Jew (Anatomy and Cell Biology), Alan Kim Johnson (Psychology), William Johnson (Microbiology), Wayne Johnson (Molecular Physiology and Biophysics), Bradley Jones (Microbiology), Barry Kasson (Pharmacology), Masataka Kawai (Anatomy and Cell Biology), Alan Kay (Biological Sciences), Shahram Khademi (Biochemistry), John Kirby (Microbiology), Toshihiro Kitamoto (Anesthesiology), Joel Kline (Internal Medicine), Al Klingelhutz (Microbiology), C. Michael Knudson (Pathology), Amnon Kohen (Chemistry), John Koland (Pharmacology), Ulla Kopp (Internal Medicine), David Kusner (Internal Medicine), Tomomi Kuwana (Pathology), Michel Ladouceur (Integrative Physiology), Kathryn Lamping (Internal Medicine), Gloria Lee (Internal Medicine), Kevin Legge (Pathology), Steven Lentz (Internal Medicine), Andrew Lidral (Orthodontics), Jack Lilien (Biological Sciences), Nancy Lill (Pharmacology), Jim (Chung-Jung) Lin (Biological Sciences), John Logsdon (Biological Sciences), David Lubaroff (Urology), Gabriele Ludewig (Occupational and Environmental Health), Vince Magnotta (Radiology), Rama Mallampalli (Internal Medicine), Robert Malone (Biological Sciences), Claudio Margulis (Chemistry), Brian Martin (Microbiology), Wendy Maury (Microbiology), Bryant McAllister (Biological Sciences), Anton McCaffrey (Microbiology), Linda McCarter (Microbiology), Paul McCray (Pediatrics), Stephen McGowan (Internal Medicine), Jeffrey Meier (Internal Medicine), John Menninger (Biological Sciences), Lou Messerle (Chemistry), Francis Miller (Internal Medicine), Steven Moore (Pathology), Craig Morita (Internal Medicine), David Moser (Psychiatry), Scott Moye-Rowley (Molecular Physiology and Biophysics), David Murhammer (Chemical and Biochemical Engineering), Jeffrey Murray (Pediatrics), Peter Nagy (Pathology), William Nauseef (Internal Medicine), Peggy Nopoulos (Psychiatry), Larry Oberley (Radiation Oncology), Sue O'Dorisio (Pediatrics), Daniel O'Leary (Psychiatry), Nicholas Pantazis (Anatomy and Cell Biology), Sergio Paradiso (Psychiatry), Shivanand Patil (Pediatrics), Jane Paulsen (Psychiatry), Stanley Perlman (Pediatrics), Robert Philibert (Psychiatry), Robert Piper (Molecular Physiology and Biophysics), Bryce Plapp (Biochemistry), Amy Poremba (Psychology), David Price (Biochemistry), Herbert Proudfit (Pharmacology), Dawn Quelle (Pharmacology), Frederick Quelle (Pharmacology), Daniel Quinn (Chemistry), Subramanian Ramaswamy (Biochemistry), Timothy Ratliff (Urology), Michael Rebagliati (Anatomy and Cell Biology), Matthew Rizzo (Neurology), Larry Robertson (Occupational and Environmental Health), Robert Robinson (Psychiatry), Jan-Uwe Rohde (Chemistry), Richard Roller (Microbiology), Paul Rothman (Internal Medicine), Peter Rubenstein (Biochemistry), Andrew Russo (Molecular Physiology and Biophysics), Alexander Sandra (Anatomy and Cell Biology), Jeffry Schabilion (Biological Sciences), Thomas Schmidt (Molecular Physiology and Biophysics), Jerald Schnoor (Civil and Environmental Engineering), Brian Schutte (Pediatrics), Deborah Segaloff (Molecular Physiology and Biophysics), Alberto Segre (Computer Science), Madeline Shea (Biochemistry), David Sheff (Pharmacology), Val Sheffield (Pediatrics), Erwin Shibata (Molecular Physiology and Biophysics), Ming-Che Shih (Biological Sciences), Curt Sigmund (Internal Medicine), Kathleen Sluka (Physical Therapy and Rehabilitation Science), Diane Slusarski (Biological Sciences), Richard Smith (Otolaryngology--Head and Neck Surgery), Jeanne Snyder (Anatomy and Cell Biology), David Soll (Biological Sciences), Arthur Spector (Biochemistry), Douglas Spitz (Radiation Oncology), Mark Stamnes (Molecular Physiology and Biophysics), Jack Stapleton (Internal Medicine), George Stauffer (Microbiology), Barbara Stay (Biological Sciences), Mark Stinski (Microbiology), Christopher Stipp (Biological Sciences), C. Martin Stoltzfus (Microbiology), Edwin Stone (Ophthalmology), Stefan Strack (Pharmacology), William Talman (Neurology), Jason Telford (Chemistry), Christie Thomas (Internal Medicine), Peter Thorne (Occupational and Environmental Health), Ingo Titze (Speech Pathology and Audiology), Robert Tomanek (Anatomy and Cell Biology), J. Bruce Tomblin (Speech Pathology and Audiology), Daniel Tranel (Neurology), Lubomir Turek (Pathology), Christopher Turner (Speech Pathology and Audiology), Ergun Uc (Neurology), Yuriy Usachev (Pharmacology), Steve Varga (Microbiology), Shaun Vecera (Psychology), Thomas Waldschmidt (Pathology), Michael Wall (Neurology), Lori Wallrath (Biochemistry), Andy (Wei-Yeh) Wang (Biological Sciences), M. Todd Washington (Biochemistry), Edward Wasserman (Psychology), Thomas Wassink (Psychiatry), Daniel Weeks (Biochemistry), George Weiner (Internal Medicine), Joshua Weiner (Biological Sciences), Jerrold Weiss (Internal Medicine), David Weiss (Microbiology), Michael Welsh (Internal Medicine), John Wemmie (Psychiatry), David Wiemer (Chemistry), Terence Williams (Anatomy and Cell Biology), Marcia Willing (Pediatrics), Mary Wilson (Internal Medicine), Marc Wold (Biochemistry), Dale Wurster (Pharmacy), Chun-Fang Wu (Biological Sciences), Yi Xing (Internal Medicine), Hai-Hui (Howard) Xue (Microbiology), Timothy Yahr (Microbiology), Charles Yeaman (Anatomy and Cell Biology), Joseph Zabner (Internal Medicine), Nicholas Zavazava (Internal Medicine) Web site: http://www.medicine.uiowa.edu/biosciences Graduate ProgramThe Biosciences Program provides graduate students the freedom to explore research in any of 14 University of Iowa biosciences research departments and programs: the Departments of Anatomy and Cell Biology, Biochemistry, Biological Sciences, Chemistry, Microbiology, Molecular Physiology and Biophysics, Pharmacology, and Speech Pathology and Audiology; and the Programs in Free Radical and Radiation Biology, Genetics, Human Toxicology, Immunology, Molecular and Cellular Biology, and Neuroscience. The program leads to a Ph.D. degree in one of the biosciences disciplines.Biosciences students enjoy the flexibility of investigating several disciplines through research rotations in the laboratories of Biosciences Program faculty members. Following completion of their first year in the program, students select a research laboratory and program affiliation and decide on a thesis project that will lead to a Ph.D. Semester hour requirements for the doctorate vary by program, but all Ph.D. degrees at Iowa require at least 72 s.h. of graduate credit. For detailed information on Graduate College policies, see the Manual of Rules and Regulations of the Graduate College. CurriculumDuring their first year in the program, students perform three 10-week research rotations in Biosciences Program departments and programs of their choice. They also take a principles course (156:201), a seminar course (156:265), and one or more electives.Biosciences students are advised on course selection, research rotations, and registration by a faculty member closely related to the student's research and academic interests. As research rotations are assigned, the faculty advisor works in consultation with the student's rotation advisors until the end of the first year. Biosciences students provide a short oral presentation following each research rotation to an audience of their primary advisor, research advisor(s), and other biosciences students. Rotation advisors provide rotation reports, and rotations are evaluated by the student's primary advisor. The student's primary advisor also confers with the student on course grades, subsequent rotations, and the student's selection of a department or program and lab for thesis research. In addition to laboratory rotations, all biosciences students take 156:201 Principles in Molecular and Cell Biology, which provides a foundation for understanding basic principles of molecular and cell biology relevant to all bioscience disciplines and teaches students how to evaluate literature critically. The Biosciences Seminar (156:265) dovetails with current seminar series offered by the Biosciences Program's departments and programs. At the beginning of each semester, biosciences students meet to choose approximately 12 seminars that they all will attend. Students choose seminars under advisement from the course instructor, who ensures that all students contribute to the selection process. During the first semester, student groups are formed and each group selects a paper from a chosen speaker to present. During the second semester, individual students select and present a paper. Overall student progress is monitored by the student's primary advisor, rotation advisor(s), and program director, who meet at the end of each semester to review the student's progress. At the end of the second semester, the primary advisor, in cooperation with the rotation advisor(s), makes a recommendation to the Biosciences Program director whether the student should continue in the Ph.D. program. University guidelines, such as maintaining a cumulative g.p.a. of 3.00 or higher, are considered, along with performance in rotations. During the second semester, Biosciences Program students meet with prospective mentors regarding thesis proposals and openings available in the mentors' laboratories. Students identify which program and laboratory they would like to join. Two weeks before the spring semester ends, students submit their choice of graduate programs and faculty mentors to the Biosciences Program office, which coordinates each student's transfer to his or her chosen program. From that point on, the department or program advises the student and evaluates his or her performance. Most participating departments and programs require that students take a comprehensive exam at the end of the second year and no later than the third year. Following successful completion of the exam, students advance to Ph.D. candidacy. REQUIRED COURSESAll Biosciences Program students must complete the following course work. Students earn at least 12 s.h. each semester.Fall Semester
Spring Semester
AdmissionThe program accepts students with a variety of backgrounds in the biological and physical sciences. Entering students must hold a baccalaureate degree from an accredited college or university and should have completed courses in biology, chemistry, physics, and calculus consistent with requirements for a baccalaureate degree in the sciences. An undergraduate g.p.a. of at least 3.00 is required.Applicants must submit their scores on the Graduate Record Examination; a combined verbal and quantitative score of 1200 and an analytical writing score of 4.5 or higher are preferred. Applicants whose first language is not English must score above 600 (paper-based), 250 (computer-based), or 100 (Internet-based) on the Test of English as a Foreign Language (TOEFL). Other indicators of academic accomplishments, such as research experience and letters of recommendation, are considered. Information about graduate training and application materials are available from the Biosciences Program office. Financial SupportGraduate students receive stipend and tuition support from The University of Iowa and other sources. Students promoted to the second year in the program receive support from their graduate departments and programs. The Biosciences Program also helps some students apply for competitive national awards for outstanding academic and research achievement.FacilitiesThe basic science and clinical departments of the Carver College of Medicine are clustered on the west campus of the University, primarily in the Bowen Science Building, Carver Biomedical Research Building, Eckstein Medical Research Building, Medical Education and Biomedical Research Facility, Medical Laboratories, and University of Iowa Hospitals and Clinics. Nearby are the Hardin Library for the Health Sciences and the Veterans Affairs Iowa City Health Care System.The Departments of Anatomy and Cell Biology, Biochemistry, Microbiology, Molecular Physiology and Biophysics, and Pharmacology are housed in the Bowen Science Building. Laboratories of clinical departments are located primarily in the Medical Laboratories and the Medical Research Center. The Eckstein Medical Research Building houses major core facilities for microscopy, image analysis, flow cytometry, protein structure, and monoclonal antibody production, as well as research laboratories for basic investigators with interdisciplinary approaches to cancer, molecular biology, genetics, and immunology. The geographic proximity of these facilities promotes interchange among clinical and basic science faculty members and students and maximizes use of the University's extensive core facilities for biomedical research. Integral to the University's research environment are the Alzheimer's Disease Research Center, Holden Comprehensive Cancer Center, Iowa Cardiovascular Center, Clinical Research Center, Cooperative Human Linkage Center, Craniofacial Anomalies Research Center, Diabetes-Endocrinology Research Center, Digestive Diseases Core Center, Iowa Child Health Research Center, University of Iowa Cystic Fibrosis Center, Iowa Specialized Center for Pulmonary Research, Lipid Research Clinic, Iowa Mental Health Clinical Research Center, Schizophrenia Research Center, Iowa Center for Gene Therapy, and Specialized Centers for Congenital Heart Disease and Hypertension and for Occupational and Immunological Lung Disease. In addition to the University's extensive facilities for research support, the Carver College of Medicine and the College of Liberal Arts and Sciences operate a variety of research support facilities. Tissue culture, autoclaving, purified water, darkrooms, counters, and a variety of general-use equipment and services are available on a shared basis. Courses
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